Program Details

Why E3?

Life has evolved numerous solutions to complex problems posed by past and present environments. In the interrelated fields of evolution and ecology, we study how and why living systems have evolved into their current forms, and how they function in complex environments. This involves both a broad comparative approach that addresses the many different ways that organisms have been able to make a living on the planet, and an interest in deep time that addresses the processes that have shaped living systems as we see them today. Scientists working in evolution and ecology have research interests spanning multiple levels of biological organization, from genetics and development to the functioning of whole organisms in complex ecological communities. This type of integrative biology is more relevant than ever, as we struggle to understand the actual range of biological diversity and how to preserve it. In particular, understanding and mitigating the effects of global climate change are among the most pressing modern concerns. The E3 REU is aimed at training highly motivated young scientists and preparing them for successful careers in the broad fields of evolution, ecology, and environmental biology.

Program Elements:

  • Multidisciplinary Research Projects
  • Weekly Professional Development Workshops
  • Networking & Social Activities
  • Supported Attendance of the Leadership Alliance National Symposium 
  • End-of-Summer Harvard-wide Poster Session

Program Information:

  • The E3 REU is committed to running a program during the summer of 2022, but it remains to be determined whether it will be remote or on campus. The 2020 and 2021 programs ran successfully in a remote format, which is ready to use again for 2022 if necessary. For now, we hope to plan for an in-person program in 2022.
  • The current planned dates for the 2022 program are Tuesday June 7 to Saturday August 13.
  • Students receive a weekly stipend of $575.
  • If the program is on campus, then travel to and from the program is covered and free on-campus housing will be provided.
  • The program provides all research and course-related expenses.


  • Students living in the student village are required to be vaccinated. Exemptions will be available for religious or medical purposes.
  • Students must provide proof of Covid immunization.
  • The first week after arrival, students will take a Covid test three (3) times. Subsequently, they will test once per week.
  • The program will cover the cost of testing.
  • It is likely that there will not be an arrival quarantine requirement – students can go into labs 24 hours after arriving/taking their first Covid test.
  • If a student tests positive, they will be required to quarantine in a dedicated space or to be sequestered in their own room (same policy and procedure that is currently in place for the college).

Eligibility Requirements:

  • Must be a Citizen or Permanent resident of the United States
  • Must be an undergraduate currently enrolled in a degree program and not graduated by the first day of the program
  • Must be 18 years of age before the first day of the program

For more information, please email Elena Kramer, or view the Summer Research Opportunities at Harvard program website